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Except where otherwise noted, this item's license is described as Wiley This is the peer reviewed version of the following article: [Cytoskeleton (Hoboken). 2017 Feb;74(2):91-103.], which has been published in final form at [https://doi.org/10.1002/cm.21347]. This article may be used for non-commercial purposes in accordance with Wiley Terms and Conditions for Use of Self-Archived Versions.